| Category | Variables | Abbreviations |
|---|---|---|
| Weighty | Body weight | BW |
| Liver weight | LW | |
| Gonad weight | GW | |
| Meristic | Dorsal fin spines count | DFSC |
| Dorsal fin soft ray count | DFSRC | |
| Anal fin spines count | AFSC | |
| Anal fin soft ray count | AFSRC | |
| Scales on the upper lateral line count | SULLC | |
| Scales on the lower lateral line count | SLLLC |